Systematic Catalog of the Culicidae
Walter Reed Biosystematics Unit

           


Subgenus  Armigeres  Theobald
Desvoidya Blanchard 1901:1046 (Dec. 6). Diatype: Culex obturbans Walker. Theobald 1903a:134 (as Desvoidea, lapsus)., (file size = 1030Kb)
Armigeres Theobald l901b:235 (July 15), l901a:98, 322 (Nov.23). Logotype: Culex obturbans Walker (Brunetti 1914:55)., (file size = 753Kb)
Blanchardiomyia Brunetti 1912:440. Diatype: Culex obturbans Walker., (file size = 0Kb)
Neosquamomyia Taylor 1914d:186. Haplotype: breinli Taylor., (file size = 1452Kb)

Important References:
Thurman 1959:76 (Thailand, Keys)., (file size = 12309Kb)
Delfinado 1966a:75 (Philippines, Keys)., (file size = 15239Kb)
Steffan 1968b:135 (Papuan Subregion, Keys)., (file size = 0Kb)

alkatirii Toma, Miyagi and Syafruddin.
Indonesia
 
1995:110 (M*, F*, P*, L*). , (file size = 0Kb)
Type-loc: Matayagan, North Sulawesi, Indonesia (NMNH).
 
apoensis  Bohart and Farner.
Philippines
 
1944:69 (M*, F). , (file size = 343Kb)
Type-loc: Sibulan River, Mt. Apo, [Davao], Mindanao, Philippines (MCZ).
Delfinado 1966a:79 (M*)., (file size = 15239Kb)
 
aureolineatus (Leicester).
Cambodia, China, India, Laos, Malaysia, Nepal, Philippines, Sri Lanka, Thailand, Vietnam
 
1908:79 (F,Desvoidya)
Type-loc: Ampang, [Selangor], Malaya (BM).
Senior-White 1927:67 and pl. VII, fig. 6 (L*)., (file size = 1013Kb)
Barraud 1934:319 (M*, F, L)., (file size = 33683Kb)
Thurman 1959:83 (M*, L)., (file size = 12309Kb)
Delfinado 1966a:80 (M*, distr.)., (file size = 15239Kb)
Tsukamoto, Miyagi and Toma 1985:153 (distr.)., (file size = Kb)
Darsie and Pradhan 1990:71 (distr.)., (file size = Kb)
Darsie 2000a 102(1): 109 (P*, tax., key)., (file size = Kb)
 
azurini  Basio.
Philippines
 
1971a:54 (M*). , (file size = 1204Kb)
Type-loc: : Mt. Makiling, Laguna, Luzon, Philippines (PNM).
 
baisasi  Stone and Thurman.
Philippines
 
1958:240 (M*, F). , (file size = 0Kb)
Type-loc: Abuyog, Leyte, Philippines (USNM).
Baisas 1935b:488 (M*, F; as kuchingensis)., (file size = 1405Kb)
Delfinado 1966a:80 (M*, F, P*, L*)., (file size = 15239Kb)
Darsie 2000a 102(1): 113 (P*; tax.; key)., (file size = Kb)
 
bhayungi Thurman and Thurman.
Thailand
 
1958:186 (F). , (file size = 232Kb)
Type-loc: Tad Muey Falls on Doi (Mountain) Sutep, Chiengmai Prov., Thailand (USNM).
 
breinli (Taylor).
Indonesia, Papua New Guinea, Solomon Islands
 
1914d:186 (F* only; Neosquamomyia). , (file size = 1452Kb)
Type-loc: Milne Bay, Papua, New Guinea (ANC).
Lee 1944c:217 (M*, F, L*)., (file size = 0Kb)
Penn 1949b:65 (P*)., (file size = 8364Kb)
Belkin 1962a:482 (M*, F, P*, L*)., (file size = 75916Kb)
Steffan 1968b:143 (M*, F, P*, L*)., (file size = 0Kb)
Rodhain and Gaxotte 1977:77 (distr.)., (file size = 256Kb)
Maffi et al. 1979:214 (distr.).
Lee, D.J., M.M. Hicks, M. Griffiths, M.L. Debenham, J.H. Bryan, R.C. Russell, M. Geary and E.N. Marks. 1988d:6 (lectotype F; type info.)., (file size = 0Kb)
 
candelabrifer  Brug.
Indonesia
 
1939:99 (M*, F, L*). , (file size = 0Kb)
Type-loc: Near Lindoe Lake, Celebes (BM).
Brug 1939, fig. 5 (as candelabris, lapsus)
 
confusus  Edwards.
Indonesia, Malaysia
 
1915c:283 (M, F). , (file size = 0Kb)
Type-loc: Kuching, Sarawak, Borneo (BM).
Bonne Wepster and Brug 1937b:529 (M, F*)., (file size = 6282Kb)
Bonne Wepster and Brug 1939:1234 (L*)., (file size = 3842Kb)
Mattingly 1971d:122 (E*)., (file size = 0Kb)
 
conjungens Edwards.
Malaysia, Singapore
 
1914c:126 (M, F). , (file size = 0Kb)
Type-loc: Kuching, Sarawak, Borneo (BM).
Mattingly and Qutubuddin 1952:92 (M*)., (file size = 0Kb)
 
denbesteni  Brug.
Indonesia, Papua New Guinea
 
1925:345 (M*, F, L*). , (file size = 161Kb)
Type-loc: Piroe, Ceram, Moluccas (BM).
Steffan 1968b:151 (M*, F, P*, L*)., (file size = 0Kb)
 
spathulatus  Brug
 
1939:101 (M*). , (file size = 0Kb)
Type-loc: Ralio Sawalit, Ceram, Moloccas (BM).
Lee 1944c:215 (syn.)., (file size = 0Kb)
Townsend 1990:136 (type info.)., (file size = 12404Kb)
 
durhami  Edwards.
Cambodia, India, Indonesia, Laos, Malaysia, Nepal, Vietnam
 
1917:206 (M, F). , (file size = 2264Kb)
Type-loc: Bukit Kutu, Malaya [Malaysia] (BM).
Thurman 1958 (1959):391 (M*; to sp. status)., (file size = 566Kb)
Thurman 1959:85 (M*, L)., (file size = 12309Kb)
Darsie and Pradhan 1990:71 (distr.). , (file size = Kb)
Darsie 2000a 102(1): 109 (P*; tax.; key)., (file size = Kb)
 
dibrugarhensis Barraud
 
1927b:539 (A; kuchingensis form). , (file size = 1447Kb)
Type-loc: Upper Assam Valley, [India] (LU).
Thurman 1959:85 (syn.)., (file size = 12309Kb)
 
javanensis Brug.
 
1939:100 (M, F, L*). , (file size = 0Kb)
Type-loc: Tangkoeban Prahoe, Java (BM).
Thurman 1959:87 (?syn.)., (file size = 12309Kb)
Ramalingam 1987:64 (syn.)., (file size = 1080Kb)
 
shillongensis Barraud
 
1927b:539 (A; kuchingensis form). , (file size = 1447Kb)
Type-loc: Assam Hills, [India] (LU).
Thurman 1959:85 (syn.)., (file size = 12309Kb)
 
ejercitoi Baisas.
Philippines
 
1935b:486 (M*, F). , (file size = 1405Kb)
Type-loc: Kolambugan, Lanao, Mindanao, Philippines (NE).
Delfinado 1966a:82 (M*, F)., (file size = 15239Kb)
 
fimbriatus Edwards.
Malaysia
 
1930a:305 (M). , (file size = 2110Kb)
Type-loc: Bettatan, near Sandakan, North Borneo, Borneo (BM).
 
foliatus Brug.
Indonesia
 
1931c:32 (M*). , (file size = 3646Kb)
Type-loc: Ranau-See, [Ranau Danau], Sumatra (BM).
 
giveni Edwards.
Singapore
 
1926a:118 (M, L). , (file size = 2849Kb)
Type-loc: Singapore (BM).
Toma and Miyagi 2002: 264 (M*)., (file size = Kb)
 
hybridus Edwards.
Malaysia, Singapore
 
1914c:126 (M, F). , (file size = 0Kb)
Type-loc: Kuching, Sarawak, Borneo (BM).
 
joloensis (Ludlow).
India, Philippines
 
1904b:236 (M, F; Desvoidea fusca var.). , (file size = 270Kb)
Type-loc: Jolo, Jolo Island, [Sulu Island], Philippines (USNM).
Stone and Thurman 1958:243 (M*; resurrected from syn. with obturbans)., (file size = 0Kb)
Bhattacharyya 2000: 63 (distr., India)., (file size = Kb)
Kaur 2003: 453 (distr., India)., (file size = Kb)
 
jugraensis  (Leicester).
Indonesia, Malaysia, Thailand
 
Type-loc: Type-loc: Ampang Jungle, Kuala Lumpur, [Selangor], Malaya (BM).
Thurman 1959:86 (M*, L)., (file size = 12309Kb)
 
kesseli Ramalingam.
Malaysia, Nepal, Thailand
 
1987:56 (M*, F*, P*, L*). , (file size = 1080Kb)
Type-loc: Ulu Gombak, Selkangor, Malaysia (USNM).
Darsie, Courtney and Pradhan 1996:133 (distr., Nepal).
 
kinabaluensis Ramalingam.
Malaysia
 
1972:460 (M*, F, P*, L*). , (file size = 536Kb)
Type-loc: Mount Kinabalu, Sabah (North Borneo), East Malaysia (USNM).
 
kuchingensis  Edwards.
Bangladesh, Cambodia, India, Laos, Malaysia, Nepal, Thailand, Vietnam
 
1915c:283 (M, F). , (file size = 0Kb)
Type-loc: Kuching Reservoir, Sarawak, Borneo (BM).
Borel 1930:186 (M*, F, L*)., (file size = 20742Kb)
Barraud 1934:317 (M*, F, L)., (file size = 33683Kb)
Edwards, in Barraud 1934:314 (tax.)., (file size = 0Kb)
Stone and Thurman 1958:240 (M*; resurrected from syn. with obturbans)., (file size = 0Kb)
Thurman 1959:86 (M*, L)., (file size = 12309Kb)
Joshi, Pradhan and Darsie 1965:138 (distr., Nepal)., (file size = 654Kb)
Ahmed 1987:187 (distr.)., (file size = Kb)
Darsie 2000a 102(1): 109 (P*; tax.; key)., (file size = Kb)
 
nongpohensis Barraud
 
1927b:539 (A; as form). , (file size = 1447Kb)
Type-loc: : Assam Hills and E. Himalayas, [India] (LU).
Thurman 1959:87 (syn.)., (file size = 12309Kb)
 
lacuum Edwards.
Papua New Guinea
 
1922d:97 (M, F). , (file size = 2717Kb)
Type-loc: Ile des Lacs (now Garove, Vitu Islands). Bismarck Archipelago (HNM) .
Hill 1925:70 (L*, P*)., (file size = 6877Kb)
Lee 1944c:219 (M*, F, L)., (file size = 0Kb)
Steffan 1968b:147 (M*, F, P*, L*; change in distr.)., (file size = 0Kb)
 
laoensis Toma and Miyagi
Laos
 
2003: 169 (M*). , (file size = Kb)
Type-loc: Thapachon, Khammouane Province, Lao PDR [Laos] (USNHM).
 
maiae Edwards.
Malaysia
 
1917:208 (M*). , (file size = 2264Kb)
Type-loc: Jungle, Pahang Road, 6 miles from Kuala Lumpur, [Selangor], Malaya (BM).
Mattingly 1971d:123 (E)., (file size = 0Kb)
 
malayi (Theobald).
China, Indonesia, Malaysia, Papua New Guinea, Philippines, Thailand
 
l901a:258 (F*; Uranotaenia).
Type-loc: Klang Jungle, Selangor Malaya [Malaysia] (BM).
Edwards 1917:207 (M*)., (file size = 2264Kb)
Brug 1924:439 (L*; as jugraensis)., (file size = 792Kb)
Bonne-Wepster and Brug 1937b:532 (M, F*)., (file size = 6282Kb)
Bonne-Wepster and Brug 1939:1234 (L*)., (file size = 3842Kb)
Knight and Chamberlain 1948:10 (P*)., (file size = 1373Kb)
Thurman 1959:88 (M*, F, P*, L*)., (file size = 12309Kb)
Steffan 1968b:141 (M*, F, P*, L*)., (file size = 0Kb)
 
fusca Theobald
 
1903a:135 (M*, F, P*, L*; Desvoidea). , (file size = 22506Kb)
Type-loc: Jugra, Malaya (BM).
Ward 1992:191 (tax.)
 
russelli  Baisas
 
1935b:490 (M*, F). , (file size = 1405Kb)
Type-loc: Masiit, Calauan, Laguna, Luzon, Philippines (NE).
Bohart 1945:47 (syn.)., (file size = 7969Kb)
 
manalangi Baisas.
Philippines
 
1935b:492 (M*, F). , (file size = 1405Kb)
Type-loc: Tungcong Manga, San Jose, Bulacan Luzon, Philippines (PBH).
Delfinado 1966a:86 (M*, F, P*, L*)., (file size = 15239Kb)
 
maximus  Edwards.
Indonesia, Malaysia, Thailand
 
1922a:292 (F). , (file size = 2704Kb)
Type-loc: Suban Ajam, [Soebanajam, Benkoelen], Sumatra (AM).
Harrison, Rattanarithikul, Peyton and Mongkolpanya 1990:224 (distr., Thailand)
Harrison, Rattanarithikul, Peyton and Mongkolpanya 1991:224 (distr,, Thailand), (file size = 0Kb)
 
milnensis Lee.
Indonesia, Papua New Guinea, Solomon Islands
 
1944c:215 (M*, F, L*). , (file size = 0Kb)
Type-loc: Milne Bay, Papua, New Guinea (CSIR).
Taylor 1914d:188 (M only, as breinli)., (file size = 1452Kb)
Penn 1949b:66 (P*)., (file size = 8364Kb)
Steffan 1968b:153 (M*, F, P*, L*)., (file size = 0Kb)
Lee et al. 1988d:123 (distr.). , (file size = 0Kb)
 
ventralis  (Walker)
 
1865 (F; Culex), (non Walker, 1860).
Type-loc: New Guinea (NMM).
Marks in Lee et al. 1988d:19 (from nomen dubium of genus Culex; syn.; type info.), (file size = 0Kb)
 
moultoni  Edwards.
Cambodia, Indonesia, Laos, Malaysia, Thailand, Vietnam
 
1914c:125 (M, F). , (file size = 0Kb)
Type-loc: Kuching, Sarawak, Borneo (BM).
Harrison, Rattanarithikul, Peyton and Mongkolpanya 1990:224 (distr., Thailand).
 
obturbans (Walker).
Indonesia, Maldives
 
1859:91 (F; Culex). , (file size = 0Kb)
Type-loc: Makassar, Celebes (NMM).
Edwards 1932a:185 (Armigeres)., (file size = 23120Kb)
Van Ty 1945:304 (biol.)., (file size = 373Kb)
Iyengar and Menon 1955:1 (distr.). , (file size = 663Kb)
Lee, D.J., M.M. Hicks, M. Griffiths, M.L. Debenham, J.H. Bryan, R.C. Russell, M. Geary and E.N. Marks. 1988d (to valid sp.; type info.)., (file size = 0Kb)
 
pallidabdomen Dong, Zhou and Gong
 
2004: ??, (file size = Kb)
 
pallithorax Dong, Zhou and Dong
China
 
2004: 125 (M*, F).
Type-loc: Xishuangbanna (elevation 780m), Yunnan Province, China (IPD).
 
papuensis Peters.
Indonesia, Papua New Guinea
 
1963a:4 (M*, F*, P*, L*). , (file size = 877Kb)
Type-loc: Maprik, Sepik District, New Guinea (ANIC).
Lee et al. 1988d:29 (distr.). , (file size = 0Kb)
 
sembeli Toma and Miyagi
Indonesia
 
2002: 257 (M*, F*, P*, L*). , (file size = Kb)
Type-loc: Langowan, North Sulawesi, Indonesia (USNM).
 
seticoxitus  Luh and Li.
China
 
1981:188 (M*, F). , (file size = 182Kb)
Type-loc: Guangxi Autonomous Region, P.R. China (ANC).
 
setifer Delfinado.
Philippines
 
1966a:87 (M*, F ). , (file size = 15239Kb)
Type-loc: : Pinigisan, Mantalingajan, Palawan, Philippines (USNM).
 
subalbatus (Coquillett)
Bangladesh, Cambodia, China, India, Indonesia, Japan, Korea, Laos, Malaysia, Myanmar (Burma), Nepal, Pakistan, Philippines, Sri Lanka, Taiwan, Taiwan, Thailand, Vietnam
 
1898:302 (F; Culex). , (file size = 58Kb)
Type-loc: Japan (USNM)
Bohart and Ingram 1946b:62 (M*, F, P*, L*).
LaCasse and Yamaguti 1950:53 (M*, F*, P*, L*).
Hara 1957:50 (F*)., (file size = 2875Kb)
Thurman 1958(1959):390 (M*; tax.)., (file size = 566Kb)
Joshi, Pradhan and Dsrsie 1965:138 (distr., Nepal)., (file size = 654Kb)
Aslamkhan 1971b:153 (distr.)., (file size = 0Kb)
Mattingly 1971d:122 (E*)., (file size = 0Kb)
Matsuo, Yoshida, and Kunou 1972: 361 (E*)., (file size = 951Kb)
Miyagi et al. 1994:21 (distr.). , (file size = Kb)
Darsie 2000a 102(1): 111 (P*; tax.; key)., (file size = Kb)
 
panalectoris  Giles
 
1901a:608 (A; Culex). , (file size = 1309Kb)
Type-loc: Amoy, China (LU).
Thurman 1959:78 (syn.)., , , , , , , , , (file size = 12309Kb)
 
ssp. chrysocorporis Hsieh and Liao.
 
1956:126 (M*, F*, L*; obturbans var.). , (file size = 426Kb)
Type-loc: Amoy, China (LU).
Harbach and Howard 2007: 41 (to ssp status), (file size = Kb)
 
theobaldi  Barraud.
India, Myanmar (Burma), Thailand
 
1934:319 (nom. nov. for apicalis Theobald 1910a, non 1908b).
Mattingly and Qutubuddin 1952:92 (M*)., (file size = 0Kb)
Thurman 1959:91 (M*, F)., (file size = 12309Kb)
Toma, Miyagi and Benjaphong 1994:11 (M*, F*, P*, L*)., (file size = 0Kb)
 
apicalis Theobald
 
1908b:291 (M, F; Leicesteria). , (file size = 1514Kb)
Type-loc: Balighai, near Puri, Orissa, India (IM).
 
yunnanensis Dong, Zhou and Dong
China
 
1995:97 (F, M*, L*, P*). , (file size = 0Kb)
Type-loc: Jinhong, Yunnan Province, China ( ).