Systematic Catalog of the Culicidae
Walter Reed Biosystematics Unit

           

Genus TOXORHYNCHITES

Important References:
Van Someren 1946b:177 (key, Ethiopian Region)., (file size = 0 Kb)
Vargas 1953b:27 (key, North and Central America)., (file size = 339 Kb)
Belkin 1968b:32 (type information). , (file size = 0 Kb)
Mattingly 1969b:13 (E*)., (file size = 0 Kb)
Berlin and Pandian 1973a:274 (Indian spp. cataloged). , (file size = 0 Kb)
Ribeiro 1992:195 (New subgenus)., (file size = 0 Kb)

Subgenus  Afrorhynchus Ribeiro
Afrorhynchus Ribeiro 1992:195. Type species Toxorhynchites lutescens Theobald., (file size = 0Kb)

aeneus  (Evans).
Cote d'Ivoire, Sierra Leone, Democratic Republic of the Congo
 
1926:102 (F, P; Megarhinus). , (file size = 0Kb)
Type-loc: Hill Station, Freetown, Sierra Leone (BM).
Wolfs 1947d:69 (P*, L*)., (file size = 343Kb)
Doucet 1960:814 (n. distr.).
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
angolensis Ribeiro
Angola
 
1992:188 (M*). , (file size = 0Kb)
Type-loc: Dundo area, northeastern Angola (IHMT).
 
brunhesi Ribeiro
Madagascar
 
2004: 250 (M*, F). , (file size = Kb)
Type-loc: Cant. Perinet, Dist. Moramanga, Prov. Tamatave, Madagascar (IRD).
 
erythrurus (Edwards).
Cote d'Ivoire, Nigeria
 
1941:30, 361 (M, F, P; Megarhinus). , (file size = 40278Kb)
Type-loc: Ibadan, [Western Provinces], Nigeria (BM).
Surtees 1958:63 (L).
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
fontenillei Ribeiro
Madagascar
 
2004: 252 (M*). , (file size = Kb)
Type-loc: Madagascar (IRD).
 
grjebinei Ribeiro
Madagascar
 
2004: 251 (M*, F). , (file size = Kb)
Type-loc: Ste. Luce, Madagascar (IRD).
 
helenae Ribeiro.
Angola
 
1992:191 (M*). , (file size = 0Kb)
Type-loc: Dundo area, northeastern Angola (IHMT).
 
kaimosi (Van Someren).
Kenya
 
1946b:184 (M, F, P, L; Megarhinus). , (file size = 0Kb)
Type-loc: Kaimosi Forest, North Kavirondo, Kenya (BM).
Corbet 1963:11 (P*).
Steffan and White 1981:118 (lectotype desig.)., (file size = 0Kb)
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
lemuriae Ribeiro
Madagascar
 
2004: 254 (F*). , (file size = Kb)
Type-loc: Prov. Fianarantsoa, Loharano, SP Manakara, Madagascar (IRD).
 
lutescens (Theobald).
Kenya, South Africa, Tanzania, Zimbabwe
 
1901a:233 (M*; Megarhinus).
Type-loc: Salisbury, Mashonaland [Southern Rhodesia] (BM).
Edwards 1941:29 (M)., (file size = 40278Kb)
Van Someren 1946b:181 (F)., (file size = 0Kb)
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
madagascarensis Ribeiro
Madagascar
 
2004: 248 (M*, F). , (file size = Kb)
Type-loc: Fort Dauphin, Madagascar (IRD).
 
nairobiensis (Van Someren).
Kenya
 
1946b:181 (M, F, L*; Megarhinus). , (file size = 0Kb)
Type-loc: City Park, Nairobi, Kenya (BM).
Steffan and White 1981:119 (lectotype desig.)., (file size = 0Kb)
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
nigeriensis Ribeiro
Nigeria
 
Type-loc: Shaki, Nigeria (BM)
2005a: 227 (M*), (file size = Kb)
 
pauliani (Doucet).
Madagascar
 
1951c:107 (M*; Megarhinus).
Type-loc: Vangaindrano, Madagascar (IRSM).
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
Ribeiro 2004: 247 (M*, key)., (file size = Kb)
 
ruwenzori (Van Someren).
Uganda
 
1948:128 (M, F, L; Megarhinus).
Type-loc: Bwamba Pass, Ruwenzori, [Toro District], Uganda (BM).
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
viridibasis (Edwards).
Burkina Faso, Cote d'Ivoire, Nigeria, Sudan, Uganda
 
1935a:133 (F; Megarhinus aeneus var.). , (file size = 1487Kb)
Type-loc: Kampala, Uganda (BM).
Edwards 1941:31 (M, F; to sp. status)., (file size = 40278Kb)
Wolfs 1947d: 66 (P*, L*)., (file size = 343Kb)
Hamon 1954a:583 (L*).
Ribeiro 1992:195.(subgeneric transfer)., (file size = 0Kb)
 
ssp. voltaicus Ribeiro
Burkina Faso
 
Type-loc: Banankeledage 18.7.54, cercle Bob-Dioulasso, Haut Volta (Burkina Faso) (Insect Collection of the Unite d''Ecologie Virale, Institut Pasteur, Paris, France)
2005a: 229 (M*), (file size = Kb)
 
wolfsi Ribeiro
Congo
 
Ribeiro 2005a :224 (M*) , (file size = Kb)
Type-loc: Coquilhatville, Congo (CMT)
 
zairensis Ribeiro
Congo
 
2005a: 227 (M*) , (file size = Kb)
Type-loc: Kwanga:Bakali-Inzia,Mapanda Fuma (CMT)
 
 
Subgenus  Ankylorhynchus Lutz
Ankylorhynchus Lutz, in Bourroul 1904:53, 65. Logotype: Megarhinus purpureus Theobald (Stone 1961:31)., (file size = 41Kb)

catharinensis (Da Costa Lima, Guitton, and Ferreira).
Brazil
 
1962:230 (M*, F, P*, L*; Ankylorhynchus). , (file size = 0Kb)
Type-loc: Brusque, Santa Catarina, Brazil (IOC).
Belkin, Schick, and Heinemann 1971:7 (type-loc info.)., (file size = 6393Kb)
 
hexacis (Martini).
Bolivia
 
1931a:2017 (F; Megarrhinus). , (file size = 816Kb)
Type-loc: Yungas von Coroico, [La Paz], Bolivia (SMT).
Belkin 1968b:33 (info. on type)., (file size = 0Kb)
Belkin 1971a:31 (info. on type)., (file size = 0Kb)
 
purpureus (Theobald).
Brazil, Uruguay
 
1901a:230 (F*; Megarhinus).
Type-loc: Manaus, Amazonas, Brazil (BM).
Dyar 1928:400 (M*, F, L*)., (file size = 0Kb)
Lane 1953:118 (M*, F, P*, L)., (file size = 0Kb)
Belkin 1968b:34 (lectotype desig.).
Belkin, Schick, and Heinemann 1971:7 (type-loc info.)., (file size = 6393Kb)
Rossi and Marinez 2003: 473 (distr., Uruguay)., (file size = Kb)
 
trichopygus (Wiedemann).
Brazil, French Guiana
 
1828:4 (A; Culex).
Type-loc: Salvador, Bahia, Brazil (SNG).
Dyar 1928:401 (F, L*)., (file size = 0Kb)
Lane 1953:115 (M*, F, P*, L)., (file size = 0Kb)
Belkin 1968b:34 (tax.)., (file size = 0Kb)
Belkin, Schick, and Heinemann 1971:7 (type-loc info.)., (file size = 6393Kb)
 
neglectus  Lutz
 
In Bourroul 1904:14 (F; Ankylorhynchus). , (file size = 41Kb)
Type-loc: Sao Paulo, Brazil (NE).
Belkin, Schick, and Heinemann 1971:7 (tax.)., (file size = 6393Kb)
 
 
Subgenus  Lynchiella Lahille
Megarhinus Robineau-Desvoidy 1827:412 (non Rafinesque 1820). Haplotype: Culex haemorrhoidalis Fabricius., (file size = 879Kb)
Lynchiella Lahille 1904:14. Diatype: Culex haemorrhoidalis Fabricius., (file size = 1431Kb)

bambusicola  (Lutz and Neiva).
Brazil, Colombia, Peru, Suriname, Venezuela
 
1913:136 (M, F, P, L; Megarhinus). , (file size = 1322Kb)
Type-loc: Petropolis [Rio de Janeiro], Brazil (IOC).
Lane 1953:131 (M*, F, P*, L)., (file size = 0Kb)
 
aldrichanus Bonne-Wepster and Bonne
 
1919(1920):179 (M; Megarhinus). , (file size = 952Kb)
Type-loc: Dam, Surinam (ITH).
Belkin 1968b:32 (lectotype desig.)., (file size = 0Kb)
 
cavalierii  Garcia and Casal.
Argentina
 
1967:436 (F, P*, L*). , (file size = 562Kb)
Type-loc: Hosteria Ruffino, Puerto Iguazu Misiones, Argentina (INM).
 
gerbergi Belkin.
Grenada
 
1977b:329 (M*, F, P*, L*). , (file size = 0Kb)
Type-loc: Industry Village, Union, St. Mark, Grenada, Southern Lesser Antilles (USNM).
 
grandiosus (Williston).
Mexico
 
1900:224 (F; Megarrhina).
Type-loc: Omilteme, Guerrero, Mexico (BM).
Lane 1953:125 (F)., (file size = 0Kb)
 
longipes Theobald
 
l901a:241 (F*; Megarhinus).
Type-loc: Mexico (BM).
 
guadeloupensis  (Dyar and Knab).
Argentina, Brazil, Colombia, Haiti, Suriname, Trinidad and Tobago, Venezuela, Guadeloupe , Montserrat , Dominica
 
1906c:254 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Guadeloupe Island, Lesser Antilles (USNM).
Lane 1953:125 (M*, F, P, L)., (file size = 0Kb)
Van Der Kuyp 1954: 65 (M, F, P, L*, E)., (file size = 4750Kb)
Belkin and Heinemann 1976a:148., (file size = 0Kb)
Belkin and Heinemann 1976b:272 (distr.)., (file size = 0Kb)
Augier et al. 2003: 99 (P*, L*; tax.)., (file size = Kb)
 
arborealis Shannon and Del Ponte
 
Type-loc: Lules, Tucuman, Argentina (NE).
1927(1928):89 (P; Megarhinus tucumanus var.). , (file size = 0Kb)
Lane 1953:125 (syn.)., (file size = 0Kb)
 
guianensis Bonne-Wepster and Bonne
 
1920:lB0 (M, F; Megarhinus; as ssp.). , (file size = 952Kb)
Type-loc: Kwakoegron (Saramacca), Surinam (ITH).
Lane 1953:125 (syn.)., (file size = 0Kb)
Belkin 1968b:33 (lectotype desig.)., (file size = 0Kb)
 
horei  Gordon and Evans
 
1922:330 (M*, F, P*, L*; Megarhinus). , (file size = 1454Kb)
Type-loc: Macapa, Amazonas, Brazil (BM).
Knight and Chamberlain 1948:18 (P*)., (file size = 1373Kb)
Lane 1951:333 (syn.)., (file size = 274Kb)
Vargas 1953a:312 (M*)., (file size = 332Kb)
Belkin 1968b: 33 (tax; lectotype desig.)., (file size = 0Kb)
 
tucumanus Brethes
 
1926a:318 (M*, F; Megarhinus). , (file size = 235Kb)
Type-loc: Concepcion, Tucuman, Argentina (BA).
Lane 1951:333 (syn.)., (file size = 274Kb)
Casal, in Belkin, Schick, and Heinemann 1968:10 (lectotype desig.)., (file size = 0Kb)
 
haemorrhoidalis (Fabricius).
Argentina, Bolivia, Brazil, Colombia, Guyana, Peru, Suriname, Venezuela, French Guiana
 
1787:364 (M; Culex). , (file size = 87Kb)
Type-loc: Cayenne,[Guyane], French Guiana (NE).
Dyar 1928:403 (M*, F)., (file size = 0Kb)
Seguy 1950:144 (M*).
Lane 1953: 119 (M, F, P, L*)., (file size = 0Kb)
Belkin 1968b:33 (info. on type)., (file size = 0Kb)
 
lynchi  Dyar and Knab
 
1906c:244 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Argentina (BA).
Lane 1939:42 (syn.)., (file size = 16792Kb)
Casal, in Belkin, Schick, and Heinemann 1968:11 (lectotype desig.)., (file size = 0Kb)
 
ssp. separatus  (Lynch Arribalzaga).
Argentina, Brazil, Paraguay
 
1891b:133 (M; Megarhina; as sp.).
Type-loc: Formosa, Argentina (BA).
Forattini and Lane 1952: 57 (P*, L*).
Lane 1951:336 (to ssp. status)., (file size = 274Kb)
Lane 1953:123 (M, F, P, L)., (file size = 0Kb)
Casal, in Belkin, Schick, and Heinemann 1968:11 (lectotype desig.)., (file size = 0Kb)
 
mariae (Bourroul).
Brazil, Trinidad and Tobago
 
1904:3 (F, L; Megarhinus). , (file size = 41Kb)
Type-loc: Ilha de Itaparica, Bahia, Brazil (NE).
Dyar 1928:404 (L*; as violaceus)., (file size = 0Kb)
Lane 1944:1B9 (M, F, P, L; from syn. with violaceus)., (file size = 1279Kb)
Belkin, Schick, and Heinemann 1971:7 (tax.)., (file size = 6393Kb)
 
iris Knab
 
1913:35 (F; Megarhinus). , (file size = 102Kb)
Type-loc: Trinidad (USNM).
Lane 1944:189 (syn.)., (file size = 1279Kb)
 
portoricensis (Roeder).
Cuba, Dominican Republic, Haiti, Jamaica, Puerto Rico, Montserrat
 
1885:337 (M; Megarrhina).
Type-loc: Puerto Rico (NE).
Lane 1953:128 (M*, F, L*)., (file size = 0Kb)
Vargas 1953a:313 (M*, L)., (file size = 332Kb)
Belkin 1968b:34 (tax.)., (file size = 0Kb)
Belkin, Heinemann, and Page 1970:233 (M*, F, P*, L*)., (file size = 38713Kb)
 
haitiensis Dyar and Knab
 
1906c:253 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: San Francisco mines, Santo Domingo [Dominican Republic] (USNM).
Belkin and Heinemann 1973:218 (type-loc. info.)., (file size = 0Kb)
 
pusillus  (Da Costa Lima).
Brazil
 
1931d:313 (M, F, L*; Megarhinus). , (file size = 788Kb)
Type-loc: Alto da Boa Vista, Tijuca, Rio de Janeiro, Brazil (IOC).
Lane 1953:130 (M, F, P*)., (file size = 0Kb)
 
rizzoi (Deus Palma and Bello Galvao).
Brazil
 
1969:3 (F, P*, L*).
Type-loc: Goianapolis, Goias, Brazil (IOC).
 
rutilus  (Coquillett).
Canada, United States
 
1896:44 (M, F; Megarhinus). , (file size = 339Kb)
Type-loc: Georgiana, Florida, United States (USNM).
Wood, Dang and Ellis 1979:91 (distr.).
Jenkins and Carpenter 1946:44 (P, L; biology)., (file size = 0Kb)
Vargas 1953a:313 (M*, L*)., (file size = 332Kb)
Vargas 1956b:33 (L*; tax.)., (file size = 197Kb)
Reinert 1970g:209 (P*)., (file size = 258Kb)
Trimble and Smith 1975:115 (biblio.)., (file size = 0Kb)
 
ssp. septentrionalis  (Dyar and Knab).
Canada, United States
 
1906c:249 (M, F; Megarhinus; as sp.). , (file size = 0Kb)
Type-loc: Woodstock, Virginia, United States (USNM).
Jenkins and Carpenter 1946:43 (P, L; biology; to ssp. status)., (file size = 0Kb)
Breland 1949a:38 (L*, E*)., (file size = 1435Kb)
Vargas 1953a:313 (M*, L*)., (file size = 332Kb)
Carpenter and LaCasse 1955:60 (M*, F*, L*)., (file size = 48079Kb)
Vargas 1956b:33 (L*; tax.)., (file size = 197Kb)
Dodge 1964:46 (L*)., (file size = 0Kb)
Parker 1977:93 (distr.)., (file size = 124Kb)
 
solstitialis  (Lutz).
Argentina, Brazil
 
1904. In Bourroul 1904:10 (M, F; Megarhinus).
Type-loc: Sao Paulo, Brazil (BM).
Lane 1953:123 (M*, F, P*,L)., (file size = 0Kb)
Vargas 1953a:312 (M*)., (file size = 332Kb)
Belkin 1968b:34 (tax.)., (file size = 0Kb)
Belkin, Schick, and Heinemann 1971:8 (lectotype desig.)., (file size = 6393Kb)
 
chrysocephalus  Theobald
 
1907:136 (M*; Megarhinus). , (file size = 38591Kb)
Type-loc: Sao Paulo, Brazil (BM).
 
mara  Anduze
 
1942a:43 (M*; Megarhinus). , (file size = 319Kb)
Type-loc: La Rivera (Rio Escalante), Zulia, Venezuela (IHC).
Lane 1944:180 (syn.)., (file size = 1279Kb)
DaCosta Lima, Guitton, and Ferreira 1962:243 (syn. suggested)., (file size = 0Kb)
Belkin, Schick, and Heinemann 1965:72 (syn.)., (file size = 0Kb)
 
theobaldi (Dyar and Knab).
Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Guyana, Mexico, Nicaragua, Panama, Suriname, Trinidad and Tobago, Uruguay, Venezuela
 
1906c:246 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Bogota (O).
Lane 1953: 129 (M*, F, P*, L)., (file size = 0Kb)
Vargas 1953a:313 (M*, L)., (file size = 332Kb)
Belkin 1968b:34 (tax.; lectotype desig.)., (file size = 0Kb)
Pecor et al. 2002: 246 (distr., Belize)., (file size = Kb)
Rossi and Marinez 2003: 473 (distr., Uruguay)., (file size = Kb)
 
ambiguus Dyar and Knab
 
1906c:246 (nom. nov. for ferox Wiedemann 1828 (in part), non Humboldt 1819; M; Megarhinus) , (file size = 0Kb)
Type-loc: Salvador, Bahia, Brazil (LU, ?NMW).
Lane 1953:129 (syn.)., (file size = 0Kb)
Belkin, 1968b:32 (info. on type)., (file size = 0Kb)
Belkin, Schick, and Heinemann 1971:8 (info. on type; type-loc. info.)., (file size = 6393Kb)
 
ferox  Wiedemann
 
1828:1 (M; Culex).
Type-loc: Salvador, Bahia, Brazil (SNG and NMW).
Belkin 1968b:32 (tax.)., (file size = 0Kb)
Belkin, Schick, and Heinemann 1971:8 (type-loc. info.; info. on type)., (file size = 6393Kb)
 
fluminensis  Peryassu
 
1908:128 (M, F, L*; Megarhinus). , (file size = 4418Kb)
Type-loc: Rio de Janeiro, Brazil (?IOC).
Da Costa Lima 1931d:312 (M*; syn.)., (file size = 788Kb)
 
hypoptes  Knab
 
1907a:50 (M; Megarhinus). , (file size = 251Kb)
Type-loc: Bluefields, Nicaragua (USNM).
Lane 1951:333 (syn.)., (file size = 274Kb)
 
moctezuma Dyar and Knab
 
1906c:251 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Rio Aranjuez, near Puntarenas, Costa Rica (USNM).
Lane 1953:129 (syn)., (file size = 0Kb)
 
moengoensis  Bonne-Wepster and Bonne
 
1923:7 (M, F, L; Megarhinus). , (file size = 295Kb)
Type-loc: Moengo, Surinam (ITH).
Lane 1939:43 (syn.)., (file size = 16792Kb)
Belkin 1968b:33 (lectotype desig.)., (file size = 0Kb)
 
posticatus  Lutz and Neiva
 
1913:139 (M, F, P; Megarhinus). , (file size = 1322Kb)
Type-loc: Petropolis, Rio de Janeiro, Brazil (IOC).
Belkin, Schick, and Heinemann 1971:8 (tax.)., (file size = 6393Kb)
 
trinidadensis  Dyar and Knab
 
1906c:252 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Trinidad (USNM).
Da Costa Lima 1931c:312 (syn.)., (file size = 878Kb)
Mattingly 1970g:34 :(L*)., (file size = 0Kb)
 
wiedemanni  Dyar and Knab
 
1906c:246 (nom. nov. for ferox Wiedemann 1828 (in part), non Humboldt 1819; Megarhinus)., (file size = 0Kb)
Lane 1953:129 (syn.)., (file size = 0Kb)
 
violaceus (Wiedemann).
Brazil
 
1820:7 (M; Culex).
Type-loc: Bahia, Brazil (NMW).
Da Costa Lima 1931c:310 (M*)., (file size = 878Kb)
Lane 1944:187 (F; tax.)., (file size = 1279Kb)
Belkin 1968b:34 (info. on type; lectotype desig.)., (file size = 0Kb)
 
 
Subgenus  Nomina Dubia 12 

aequatorianus  Levi-Castillo.
Ecuador
 
1953h:72(M*, F, P*, L*; Toxorhynchites). , (file size = 460Kb)
Type-loc: Hacienda Pichilingues Los Rios, Ecuador (CEIE).
Stone, Knight, and Starcke 1959:65 (nom. dub.), (file size = 27461Kb)
 
 
Subgenus  Toxorhynchites Theobald
Toxorhynchites Theobald 1901b:234 (July 15) and 1901a:244 (Nov. 23). Haplotype: brevipalpis Theobald (Int. Comm. Zool. Nomencl. 1959, Opinion 548)., (file size = 753Kb)
Worcesteria Banks 1906a:779. Orthotype: grata Banks., (file size = 750Kb)
Teromyia Leicester 1908:49. Logotype: acaudata Leicester (Brunetti 1914:72)., (file size = 0Kb)

acaudatus  (Leicester).
Indonesia, Malaysia, Singapore
 
1908:49 (M, F; Teromyia).
Type-loc: Singapore (BM).
Edwards 1926a:114 (P*, L)., (file size = 2849Kb)
Steffan and White 1981:116 (lectotype desig.)., (file size = 0Kb)
 
albipes (Edwards).
Cambodia, India, Laos, Thailand, Vietnam
 
1922a:287 (M, F; Megarhinus). , (file size = 2704Kb)
Type-loc: Simla, [East Punjab], India (BM).
Borel 1930:116 (F*, L*)., (file size = 20742Kb)
Barraud 1934:16 (M, F, P*, L*)., (file size = 33683Kb)
 
amboinensis  (Doleschall).
Fiji, Indonesia, Malaysia, Papua New Guinea, Philippines, Thailand, United States, American Samoa
 
1857:381 (A; Culex). , (file size = 0Kb)
Type-loc: Amboina Island, Moluccas (NMW).
Knight and Chamberlain 1948:11 (P*)., (file size = 1373Kb)
Steffan 1968a:150 (M*, F, P*, L*; syn.)., (file size = 0Kb)
Mattingly 1973b:197 (E*)., (file size = 0Kb)
Ramalingam and Belkin 1976:194 (distr.)., (file size = 0Kb)
Lee et al. 1988c:72 (distr.). , (file size = 0Kb)
 
argenteotarsis Ludlow
 
1906d:367 (F). , (file size = 160Kb)
Type-loc: Margosatubig, [Zamboanga], Mindanao, Philippines (USNM).
Bohart 1945:23 (syn.)., (file size = 7969Kb)
 
grata Banks
 
1906a:780 (M*, F*; Worcesteria). , (file size = 750Kb)
Type-loc: Bago, Haci enda "Louisiana", Mailum; Negros Occidental, and Cebu, Philippines (USNM).
Bohart 1945:23 (syn.)., (file size = 7969Kb)
 
immisericors  Walker
 
1859:90 (M; Megarhina). , (file size = 0Kb)
Type-loc: Makessar, Celebes (BM).
Steffan and White 1981:118 (lectotype desig.)., (file size = 0Kb)
 
lewaldii Ludlow
 
1904b:233 (M; Megarhinus). , (file size = 270Kb)
Type-loc: Salog, Gui maras Island, [Iloilo], Philippines (USNM).
Blanchard 1905:625 (as waldi; emend.)., (file size = 51185Kb)
 
subulifer Doleschall
 
1857:382 (F; Culex; as sp.). , (file size = 0Kb)
Type-loc: Amboina Island, [Moluccas] (NMW).
Knight, Bohart, and Bohart 1944: 12 (M; to ssp. status)., (file size = 4082Kb)
Bonne-Wepster 1954:11 (A)., (file size = 0Kb)
 
angustiplatus Evenhuis and Steffan.
Malaysia
 
1986:548 (M*, F, P*, L*). , (file size = 0Kb)
Type-loc: Pacific Tin, Berjuntai, Batang, Selangor, Malaysia.
 
ater (Daniels).
Malaysia
 
1908:265 (M, F, P; Teromyia). , (file size = 447Kb)
Type-loc: Pahang, Malaya (NE).
 
auranticauda Lane.
Indonesia
 
1992:301 (F*, P*, L*). , (file size = 0Kb)
Type-loc: Dumoga-Bone National Park, North Sulawesi Province, Indonesia (BMNH).
 
aurifluus (Edwards).
Indonesia, Taiwan
 
1921a:631 (M, F; Megarhinus). , (file size = 268Kb)
Type-loc: Several localities on Formosa (DEI).
Brug 1932:73 (P*)., (file size = 757Kb)
Brug 1939:92 (?F)., (file size = 0Kb)
Lien 1965:9 (M*, F*, P*, L*)., (file size = 1606Kb)
 
formosensis  Ogasawara
 
1939:242 (M; as var.).
Type-loc: Ikenohate (Shinchiku), Formosa [Chiehtuan, Loswei Village, Tatung Township, Ilan Hsien] (LU).
Lien 1962:617 (syn.)., (file size = 2678Kb)
 
auripes (Edwards).
Malaysia
 
1935b:579 (F; Megarhinus). , (file size = 0Kb)
Type-loc: Mt. Dulit, Sarawak, Borneo (BM).
 
barbipes Edwards.
Kenya, Liberia, Uganda
 
1913b:47 (M). , (file size = 1048Kb)
Type-loc: Mpanga Forest, Toro District, Uganda (BM).
Edwards 1941:28 (M*)., (file size = 40278Kb)
Van Someren 1946b:179 (M, F, P, L)., (file size = 0Kb)
 
bengalensis  Roosenberg and Evenhuis.
Bangladesh
 
1985:36 (M*, P*).
Type-loc: Chunaraghat, Sylhet District, Bangladesh (USNM).
 
bickleyi  Thurman.
Thailand
 
1959:14 (M). , (file size = 12309Kb)
Type-loc: Doi Sutep, Chiengmai Province, Thailand (USNM).
 
brevipalpis  Theobald.
Angola, Kenya, Madagascar, Mozambique, South Africa, Tanzania, United States, Zambia, Zimbabwe, Democratic Republic of the Congo
 
1901a:245 (F*).
Type-loc: [Durban], Natal (BM).
White 1977c:228 (request to ICZN for conservastion of name under plenary powers).
Edwards 1941: 26 (M, F), 360 (P)., (file size = 40278Kb)
Muspratt 1951:355 (L*, E*; biology)., (file size = 0Kb)
Doucet 1951a-82 (L*)., (file size = 0Kb)
Hopkins 1952:35 (L*)., (file size = 0Kb)
Vargas 1953a:312 (M*)., (file size = 332Kb)
Worth and De Meillon 1960:234 (distr.)., (file size = 1635Kb)
Belkin 1962a:530 (M*, F, P*, L*)., (file size = 75916Kb)
Da Cunha Ramos and ribeiro 1975:114 (distr.)., (file size = 3412Kb)
Steffan and White 1981:117 (lectotype desig.)., (file size = 0Kb)
 
marshallii Theobald
 
1903a:121 (M*). , (file size = 22506Kb)
Type-loc: Salisbury, Mashonaland, [Zimbabwe] (BM).
 
schultzei Enderlein
 
1931:123 (F). , (file size = 208Kb)
Type-loc: Mongumba "am Uban Fluss
Edwards 1941:27 (syn.)., (file size = 40278Kb)
 
tessmanni  Enderlein
 
1931:124 (F). , (file size = 208Kb)
Type-loc: Alen Benito, Uam Region, [Rio Muni] (ZM).
Edwards 1941:27 (syn.)., (file size = 40278Kb)
 
ssp. abyssinicus  Ribeiro.
Ethiopia
 
1991:43 (M*). , (file size = 1825Kb)
Type-loc: Keffa Prov., Ethiopia (BM).
 
camaronis Ribeiro.
Cameroon
 
Type-loc: Ototomo, Cameroon (SSC).
1991:50 (M*, F)., (file size = 1825Kb)
 
capelai Ribeiro.
Sao Tome and Principe
 
1992:135 (M*, F). , (file size = 0Kb)
Type-loc: Sao Tome (FCLM).
 
christophi  (Portschinsky).
Japan, Korea, Russia
 
1884:122 (F; Megarhina).
Type-loc: Amur, U.S.S.R. (LU).
Monchadskii 1951:103 (L*)., (file size = 1542Kb)
Chow 1973:43 (distr.)., (file size = 0Kb)
Gutsevich et al 1974:120 (F*, L*).
Shamray and Gutsevich 1974:427 (M*)., (file size = 167Kb)
 
changbaiensis  Su and Wang.
China
 
1981:327 (M*, F, P*, L*). , (file size = 340Kb)
Type-loc: Huang-song-pu forest area, Antu Xian, Jilin Prov., China (BMC).
Danilov 1987:154 (syn).
 
coeruleus (Brug).
Indonesia
 
1934a:501 (M, F; Megarhinus). , (file size = 122Kb)
Type-loc: Batang Palupuh, [West Coast], Sumatra (BM).
Steffan and White 1981:117 (lectotype desig.)., (file size = 0Kb)
 
dundo Ribeiro.
Angola
 
Type-loc: Dundo, Angola (IHMT).
 
edwardsi  (Barraud).
India
 
1924d:999 (M,F; Megarhinus). , (file size = 519Kb)
Type-loc: Krol Mountain, near Solon, Western Himalayas, India (BM).
Barraud 1934:23 (M, F, P*, L*)., (file size = 33683Kb)
Steffan and White 1981:117 (lectotype desig.)., (file size = 0Kb)
 
evansae  (Edwards).
Cameroon, Sierra Leone
 
1936:53 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Freetown, Sierra Leone (BM).
 
funestus (Leicester).
Malaysia
 
1908:58 (M; Teromyia).
Type-loc: Kuala Lumpur, [Selangor], Malaya (BM).
 
gigantulus  (Dyar and Shannon).
Philippines
 
1925a:67 (M; Megarhinus). , (file size = 0Kb)
Type-loc: Limay, Bataan, Luzon, Phillippines (USNM).
Bohart 1945:23 (M)., (file size = 7969Kb)
Berlin 1969b:21 (M*, F, P*, L*)., (file size = 408Kb)
 
gravelyi  (Edwards).
India, Thailand
 
1921c:73 (M; Megarhinus). , (file size = 1508Kb)
Type-loc: Pashok, [West Bengal], India (IM).
Barraud 1934:20 (M*, F, P*, L*)., (file size = 33683Kb)
 
indicus Evenhuis and Steffan.
Malaysia
 
1986: 558 (M*, F). , (file size = 0Kb)
Type-loc: Kuching, Sarawak, Malaysia (BM).
 
inornatus (Walker).
Australia, Fiji, Papua New Guinea
 
1865:102 (M; Megarhina).
Type-loc: New Guinea (BM).
Hill 1925: 67 (P*, L*)., (file size = 6877Kb)
Laird 1947:455 (L*; biology)., (file size = 2018Kb)
Belkin 1962a:533 (M*, F, P*, L*)., (file size = 75916Kb)
Steffan and White 1981:118 (lectotype desig.)., (file size = 0Kb)
 
ssp. albitarsis  (Brug).
Indonesia
 
1939:91 (M; Megarhinus; as var.). , (file size = 0Kb)
Type-loc: Kalawara, Celebes (RM) [to ssp. status].
 
kempi (Edwards).
Cambodia, India, Indonesia, Laos, Philippines, Vietnam
 
1921c:72 (M, F; Megarhinus). , (file size = 1508Kb)
Type-loc: Talewadi, North Kanara, [Bombay], India (IM).
Borel 1930:119 (M*, F*, L*)., (file size = 20742Kb)
Barraud 1934: 18 (M*, F*, P*, L*)., (file size = 33683Kb)
Tsukamoto et al. 1985:157 (distr.). , (file size = Kb)
 
klossi (Edwards).
India, Malaysia
 
1921c:72 (M; Megarhinus). , (file size = 1508Kb)
Type-loc: Kedah Peak (Gunong Jerai), [Kedah], Malaya (BM).
 
leicesteri  Theobald.
Malaysia, Philippines, Singapore, Thailand
 
1904c:36 (M, F).
Type-loc: Kuala Lumpur, [Selangor], Malaya (BM).
Edwards 1926:114 (P*, L).
Steffan and White 1981:118 (lectotype desig.)., (file size = 0Kb)
Tsukamoto et al. 1985:157 (distr.). , (file size = Kb)
 
lewisi  Ribeiro.
Sierra Leone
 
1991:53 (M*, F). , (file size = 1825Kb)
Type-loc: Farangbaia, Sierra Leone (BM).
 
macaensis Ribeiro.
China
 
1997:213 ( M*, F). , (file size = 503Kb)
Type-loc: Paque Siac Pai Van, Ilha de Coloane, Macau, China.
 
magnificus (Leicester).
Malaysia, Singapore, Thailand
 
1908:54 (M, F; Teromyia).
Type-loc: Malaya (BM).
Edwards 1926a: 114 (P*)., (file size = 2849Kb)
Steffan and White 1981:119 (lectotype desig.)., (file size = 0Kb)
 
manicatus (Edwards).
Taiwan
 
1921a:630 (F; Megarhinus). , (file size = 268Kb)
Type-loc: Toa Tsui Kutsu, Formosa (DEI).
Lien 1965:2 (M*, F*, P*, L*)., (file size = 1606Kb)
 
ssp. yaeyamae  Bohart.
Japan
 
1956:29 (M, F, P, L). , (file size = 401Kb)
Type-loc: Nakara River, Iriomote-shima, Ryukyu-Retto (USNM).
Tanaka, Saugstad and Mizusawa 1975:225 (distr.; tax)., (file size = 0Kb)
Tanaka, Mizusawa and Saugstad 1979:496 (to subspecies)., (file size = 0Kb)
 
manopi  Thurman.
Thailand
 
1959:16 (M). , (file size = 12309Kb)
Type-loc: Doi Sutep, Chiengmai Province, Thailand (USNM).
 
metallicus Leicester.
India, Indonesia, Malaysia
 
1904. In Theobald 1904c:37 (M, F).
Type-loc: Kuala Lumpur, [Selangor], Malaya (BM).
Steffan and White 1981:119 (lectotype desig.)., (file size = 0Kb)
 
minimus  (Theobald).
India, Indonesia, Philippines, Sri Lanka
 
1905h:237 (M*; Megarhinus). , (file size = 1010Kb)
Type-loc: Yatiyantota, Ceylon (BM).
Barraud 1934:22 (M, F, P*, L*)., (file size = 33683Kb)
Tsukamoto et al. 1985:157 (distr.). , (file size = Kb)
 
nepenthicola Steffan and Evenhjuis.
Papua New Guinea
 
1982:2 (M*, F*, P*, L*). , (file size = 0Kb)
Type-loc: near Olosobiip, Upper fly River, Western district, Paupa New Guinea (BBM).
 
nepenthis (Dyar and Shannon).
Philippines
 
1925a:66 (F; Megarhinus). , (file size = 0Kb)
Type-loc: Los Banos, Laguna, Luzon, Philippines (USNM).
 
nigerrmus Dong, Zhou and Gong
 
2004: ??, (file size = Kb)
 
nigripes (Edwards).
Malaysia
 
1935b:580 (M, F; Megarhinus). , (file size = 0Kb)
Type-loc: Mt. Dulit, Sarawak, Borneo (BM).
Steffan and White 1981:119 (lectotype desig.)., (file size = 0Kb)
 
okinawensis Toma, Miyagi and Tanaka.
Japan
 
1990: 345 (M*, F, P*, L*). , (file size = 976Kb)
Type-loc: Ada, Okinawajima, Japan (HUJ).
 
pendleburyi (Edwards).
Malaysia
 
1930a:305 (F; Megarhinus). , (file size = 2110Kb)
Type-loc: Kenokok, Mt. Kinabalu, North Borneo, Borneo (BM).
 
phytophagus Theobald.
Cameroon, Cote d'Ivoire, Ghana, Liberia, Sierra Leone, Democratic Republic of the Congo
 
1909:1 (F; as phytophygus). , (file size = 859Kb)
Type-loc: Obuasi, Ashanti, Gold Coast (BM).
Edwards 1912c:3 (emend.)., (file size = 0Kb)
Edwards 1941:28 (M, F)., (file size = 40278Kb)
Wolfs 1947d:63 (P*, L*)., (file size = 343Kb)
 
quasiferox (Leicester).
Indonesia, Malaysia, Singapore
 
1908:51 (M, F; Teromyia).
Type-loc: Jugra, Malaya, and Singapore (BM).
Edwards 1926a:114 (P*, L)., (file size = 2849Kb)
Steffan and White 1981:120 (lectotype desig.)., (file size = 0Kb)
 
javaensis Theobald
 
1911d:233 (F). , (file size = 347Kb)
Type-loc: Java (AM).
 
rajah Tsukamoto.
Malaysia
 
1989: 221 (M, F, P*, L*). , (file size = 999Kb)
Type-loc: Mt. Kinabalu, Sabah, Malaysia (NMST).
 
ramalingami  Evenhuis and Steffan.
Brunei
 
1986:566 (M*, F, P*, L*). , (file size = 0Kb)
Type-loc: Bukit Puan forest Reserve, Belait Dist., Brunei (USNM).
 
raris (Leicester).
Malaysia
 
1908:56 (M, L; Teromyia).
Type-loc: Ulu Klang, [Selangor], Malaya (NE).
 
rickenbachi Ribeiro.
Cameroon
 
1991:54 (M*, F).
Type-loc: Elouden, Cameroon (SSC)(P)
 
rodhaini Ribeiro.
Cote d'Ivoire
 
1991:57 (F*).
Type-loc: Tai Forest, Ivory Coast (PIP).
 
speciosus  (Skuse).
Australia, Fiji, Indonesia, Singapore
 
1889:1722 (M*; Megarrhina). , (file size = 3639Kb)
Type-loc: Port Denison, Queensland, Australia (ANIC).
Colledge 1911:121 (M, F*, P, L*, E*)., (file size = 1066Kb)
Lee 1944a: 14 (L*)., (file size = 4911Kb)
Lee, Hicks, Debenham, Griffiths, Bryan and Marks 1988c:90 (type info.; distr.)., (file size = 0Kb)
 
splendenroides Dong, Zhou and Gong
 
2004: ??, (file size = Kb)
 
splendens (Wiedemann).
Australia, Bangladesh, China, Fiji, India, Indonesia, Malaysia, Myanmar (Burma), Nepal, Papua New Guinea, Philippines, Sri Lanka, Thailand, United States
 
1819:2 (M; Culex). , (file size = 131Kb)
Type-loc: Java (ZMC and NMW).
Barraud 1934: 24 (M*, F*, P*, L*, E)., (file size = 33683Kb)
Newkirk 1947:522 (biology)., (file size = 488Kb)
Bonne-Wepster 1954:10 (M*, F*, L*)., (file size = 0Kb)
Belkin 1962a:532 (M*, F, P*, L*)., (file size = 75916Kb)
Ameen and Talukdar 1974a:201 (P*)., (file size = 0Kb)
Bhat 1976:266 (chromosomes*).
Ahmed 1987:187 (distr.). , (file size = Kb)
Lee et al. 1988c:102 (distr.). , (file size = 0Kb)
Darsie and Pradhan 1990:71 (distr.). , (file size = Kb)
 
gilesii  Theobald
 
1901a:227 (M, F*; Megarhinus).
Type-loc: Upper Burma, and Sikkim, India (BM).
Steffan and White 1981:118 (lectotype desig.)., (file size = 0Kb)
 
regius  Tennent
 
1859:268 (A; Culex).
Type-loc: Kandy, [Central], Ceylon (NE).
 
sikkimensis Giles
 
1901a:604 (F; Megarhina). , (file size = 1309Kb)
Type-loc: Sikkim, India (BM).
Barraud 1934:24 (syn.)., (file size = 33683Kb)
 
sumatranus  (Brug).
Indonesia
 
1939:93 (F; Megarhinus). , (file size = 0Kb)
Type-loc: Benkoelen, Sumatra (BM).
 
sunthorni Thurman.
Thailand
 
Type-loc: Doi Sutep, Chiengmai Province, Thailand (USNM).
1959:19 (F)., (file size = 12309Kb)
 
towadensis (Matsumura).
Japan
 
1916:445 (M*; Megarrhina).
Type-loc: Towada, Honshu, Japan (LU).
LaCasse and Yamaguti 1950:38 (M, F*).
Ishimura 1952:12 (P*, L*)., (file size = 483Kb)
Asanuma and Nakagawa 1954b:B2 (P*)., (file size = 423Kb)
Hara 1957:48 (F*)., (file size = 2875Kb)